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Base Complementarity

Definition

The specific pairing rules that govern double-stranded DNA: adenine pairs with thymine (2 hydrogen bonds) and guanine pairs with cytosine (3 hydrogen bonds). Any other combination is geometrically incompatible with the double helix. This rule is why knowing one strand's sequence immediately specifies the other strand's sequence.

A pairs with
Thymine, via 2 hydrogen bonds
G pairs with
Cytosine, via 3 hydrogen bonds
Basis of
Double helix geometry
Practical consequence
One strand's sequence fixes the other

Common questions

Why does base complementarity matter for STR profiling specifically?+

PCR amplification and probe or primer design both depend on knowing the exact complementary sequence flanking a repeat region, so primers can bind and copy only the target locus. Without reliable complementary pairing, the targeted amplification that STR typing depends on would not work.

What happens when a base pairs with a non-complementary partner?+

A mismatch like that is geometrically strained and destabilises the helix at that position, which is why DNA polymerases have proofreading mechanisms to detect and correct such errors during replication. Occasional uncorrected mismatches are one source of natural mutation.

Related terms

Antiparallel Orientation
The two strands of the DNA double helix run in opposite directions: one strand reads 5' to 3', its complement reads 3'...
Denaturation
The separation of double-stranded DNA into two single strands by disrupting the hydrogen bonds between base pairs. In PCR, denaturation is achieved...
Deoxyribonucleotide
The monomer unit of DNA, consisting of a deoxyribose sugar, a phosphate group, and one of four nitrogenous bases (adenine, thymine, guanine,...
GC Content
The proportion of base pairs in a DNA molecule that are guanine-cytosine. Because G-C pairs form three hydrogen bonds (versus two for...
Phosphodiester Backbone
The repeating sugar-phosphate chain that forms the structural scaffold of each DNA strand. The backbone is highly charged and hydrophilic, faces outward...

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