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Epitheca and Hypotheca

Definition

The two halves of the frustule. The epitheca (larger) fits over the hypotheca (smaller) like a lid over a box. During cell division each daughter cell retains one half and builds a new opposing half, so average cell size decreases over generations until sexual reproduction resets it.

Epitheca
Larger half, fits over the hypotheca
Hypotheca
Smaller half, fits inside the epitheca
Division effect
Average cell size shrinks across generations
Reset mechanism
Sexual reproduction

Common questions

Why does diatom size decline matter for forensic diatom analysis?+

Since a population's size range shifts predictably between reproductive resets, size-frequency distribution in a water or sediment sample can support comparing a questioned sample's diatom assemblage against a reference site, alongside species composition.

How does the epitheca/hypotheca structure relate to species identification?+

The frustule's valve shape, ornamentation, and the epitheca-hypotheca fit are part of the morphological features used to key out diatom species under the microscope, independent of the size-reduction cycle itself.

Related terms

Assemblage
The full community of diatom taxa present in a sample. The relative frequencies and absolute counts of species in an assemblage reflect...
Frustule
The silica cell wall of a diatom, composed of two overlapping valves (epitheca and hypotheca). Frustules are taxonomically distinctive, chemically resistant to...
Raphe
A longitudinal slit or canal in the valve of certain pennate diatoms that allows cytoplasmic streaming and active gliding movement across surfaces....
Sporopollenin vs Biogenic Silica
Pollen and spore walls are made of sporopollenin (a highly resistant biopolymer). Diatom frustules are made of biogenic opaline silica (SiO2·nH2O), a...
Valve and Girdle Band
The valve is the flat face of each frustule half, carrying the species-specific pore patterns used for identification. The girdle band is...

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